Assuming everything is in well maintained proper working condition, the biggest mileage factor while towing is speed (specifically, aerodynamics). Slowing down to the lowest safe speed will get you the most mileage benefit.
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Curious what the KO2’***** was. Use to run them on my old Land Cruiser which got 11mpg regardless of anything done to improve mileage . Running BFG trail Terrain on the Expedition and really liking them. Best average hit from stock tires might be just shy of 1mpg at highway speeds. Heavy tow pack 20-23mpg long trip averages nakid no trailer is pretty standard. Guessing the KO’s are a bigger hit?